Enzyme Nomenclature. Recommendations
EC 2.3.1.1 amino-acid N-acetyltransferase
EC 2.3.1.2 imidazole N-acetyltransferase
EC 2.3.1.3 glucosamine N-acetyltransferase
EC 2.3.1.4 glucosamine-phosphate N-acetyltransferase
EC 2.3.1.5 arylamine N-acetyltransferase
EC 2.3.1.6 choline O-acetyltransferase
EC 2.3.1.7 carnitine O-acetyltransferase
EC 2.3.1.8 phosphate acetyltransferase
EC 2.3.1.9 acetyl-CoA C-acetyltransferase
EC 2.3.1.10 hydrogen-sulfide S-acetyltransferase
EC 2.3.1.11 thioethanolamine S-acetyltransferase
EC 2.3.1.12 dihydrolipoyllysine-residue acetyltransferase
EC 2.3.1.13 glycine N-acyltransferase
EC 2.3.1.14 glutamine N-phenylacetyltransferase
EC 2.3.1.15 glycerol-3-phosphate 1-O-acyltransferase
EC 2.3.1.16 acetyl-CoA C-acyltransferase
EC 2.3.1.17 aspartate N-acetyltransferase
EC 2.3.1.18 galactoside O-acetyltransferase
EC 2.3.1.19 phosphate butyryltransferase
EC 2.3.1.20 diacylglycerol O-acyltransferase
EC 2.3.1.21 carnitine O-palmitoyltransferase
EC 2.3.1.22 2-acylglycerol O-acyltransferase
EC 2.3.1.23 1-acylglycerophosphocholine O-acyltransferase
EC 2.3.1.24 sphingosine N-acyltransferase
EC 2.3.1.25 plasmalogen synthase
EC 2.3.1.26 sterol O-acyltransferase
EC 2.3.1.27 cortisol O-acetyltransferase
EC 2.3.1.28 chloramphenicol O-acetyltransferase
EC 2.3.1.29 glycine C-acetyltransferase
EC 2.3.1.30 serine O-acetyltransferase
EC 2.3.1.31 homoserine O-acetyltransferase
EC 2.3.1.32 lysine N-acetyltransferase
EC 2.3.1.33 histidine N-acetyltransferase
EC 2.3.1.34 D-tryptophan N-acetyltransferase
EC 2.3.1.35 glutamate N-acetyltransferase
EC 2.3.1.36 D-amino-acid N-acetyltransferase
EC 2.3.1.37 5-aminolevulinate synthase
EC 2.3.1.38 [acyl-carrier-protein] S-acetyltransferase
EC 2.3.1.39 [acyl-carrier-protein] S-malonyltransferase
EC 2.3.1.40 acyl-[acyl-carrier-protein]phospholipid O-acyltransferase
EC 2.3.1.41 β-ketoacyl-acyl-carrier-protein synthase I
EC 2.3.1.42 glycerone-phosphate O-acyltransferase
EC 2.3.1.43 phosphatidylcholinesterol O-acyltransferase
EC 2.3.1.44 N-acetylneuraminate 4-O-acetyltransferase
EC 2.3.1.45 N-acetylneuraminate 7-O(or 9-O)-acetyltransferase
EC 2.3.1.46 homoserine O-succinyltransferase
EC 2.3.1.47 8-amino-7-oxononanoate synthase
EC 2.3.1.48 histone acetyltransferase
EC 2.3.1.49 deacetyl-[citrate-(pro-3S)-lyase] S-acetyltransferase
EC 2.3.1.50 serine C-palmitoyltransferase
EC 2.3.1.51 1-acylglycerol-3-phosphate O-acyltransferase
EC 2.3.1.52 2-acylglycerol-3-phosphate O-acyltransferase
EC 2.3.1.53 phenylalanine N-acetyltransferase
EC 2.3.1.54 formate C-acetyltransferase
EC 2.3.1.55 now EC 2.3.1.82
EC 2.3.1.56 aromatic-hydroxylamine O-acetyltransferase
EC 2.3.1.57 diamine N-acetyltransferase
EC 2.3.1.58 2,3-diaminopropionate N-oxalyltransferase
EC 2.3.1.59 gentamicin 2'-N-acetyltransferase
EC 2.3.1.60 gentamicin 3'-N-acetyltransferase
EC 2.3.1.61 dihydrolipoyllysine-residue succinyltransferase
EC 2.3.1.62 2-acylglycerophosphocholine O-acyltransferase
EC 2.3.1.63 1-alkylglycerophosphocholine O-acyltransferase
EC 2.3.1.64 agmatine N4-coumaroyltransferase
EC 2.3.1.65 bile acid-CoA:amino acid N-acyltransferase
EC 2.3.1.66 leucine N-acetyltransferase
EC 2.3.1.67 1-alkylglycerophosphocholine O-acetyltransferase
EC 2.3.1.68 glutamine N-acyltransferase
EC 2.3.1.69 monoterpenol O-acetyltransferase
EC 2.3.1.70 deleted
EC 2.3.1.71 glycine N-benzoyltransferase
EC 2.3.1.72 indoleacetylglucoseinositol O-acyltransferase
EC 2.3.1.73 diacylglycerolsterol O-acyltransferase
EC 2.3.1.74 chalcone synthase
EC 2.3.1.75 long-chain-alcohol O-fatty-acyltransferase
EC 2.3.1.76 retinol O-fatty-acyltransferase
EC 2.3.1.77 triacylglycerolsterol O-acyltransferase
EC 2.3.1.78 heparan-α-glucosaminide N-acetyltransferase
EC 2.3.1.79 maltose O-acetyltransferase
EC 2.3.1.80 cysteine-S-conjugate N-acetyltransferase
EC 2.3.1.81 aminoglycoside 3-N-acetyltransferase
EC 2.3.1.82 aminoglycoside 6'-N-acetyltransferase
EC 2.3.1.83 phosphatidylcholinedolichol O-acyltransferase
EC 2.3.1.84 alcohol O-acetyltransferase
EC 2.3.1.85 fatty-acid synthase
EC 2.3.1.86 fatty-acyl-CoA synthase
EC 2.3.1.87 aralkylamine N-acetyltransferase
EC 2.3.1.88 now covered by EC 2.3.1.254, EC 2.3.1.255, EC 2.3.1.256, EC 2.3.1.257, EC 2.3.1.258, EC 2.3.1.259
EC 2.3.1.89 tetrahydrodipicolinate N-acetyltransferase
EC 2.3.1.90 β-glucogallin O-galloyltransferase
EC 2.3.1.91 sinapoylglucosecholine O-sinapoyltransferase
EC 2.3.1.92 sinapoylglucosemalate O-sinapoyltransferase
EC 2.3.1.93 13-hydroxylupinine O-tigloyltransferase
EC 2.3.1.94 6-deoxyerythronolide-B synthase
EC 2.3.1.95 trihydroxystilbene synthase
EC 2.3.1.96 deleted
EC 2.3.1.97 glycylpeptide N-tetradecanoyltransferase
EC 2.3.1.98 chlorogenateglucarate O-hydroxycinnamoyltransferase
EC 2.3.1.99 quinate O-hydroxycinnamoyltransferase
EC 2.3.1.100 [myelin-proteolipid] O-palmitoyltransferase
EC 2.3.1.101 formylmethanofurantetrahydromethanopterin N-formyltransferase
EC 2.3.1.102 N6-hydroxylysine O-acetyltransferase
EC 2.3.1.103 sinapoylglucosesinapoylglucose O-sinapoyltransferase
EC 2.3.1.104 deleted, now covered by EC 2.3.1.25
EC 2.3.1.105 alkylglycerophosphate 2-O-acetyltransferase
EC 2.3.1.106 tartronate O-hydroxycinnamoyltransferase
EC 2.3.1.107 deacetylvindoline O-acetyltransferase
EC 2.3.1.108 α-tubulin N-acetyltransferase
EC 2.3.1.109 arginine N-succinyltransferase
EC 2.3.1.110 tyramine N-feruloyltransferase
EC 2.3.1.111 mycocerosate synthase
EC 2.3.1.112 D-tryptophan N-malonyltransferase
EC 2.3.1.113 anthranilate N-malonyltransferase
EC 2.3.1.114 3,4-dichloroaniline N-malonyltransferase
EC 2.3.1.115 isoflavone-7-O-β-glucoside 6"-O-malonyltransferase
EC 2.3.1.116 flavonol-3-O-β-glucoside O-malonyltransferase
EC 2.3.1.117 2,3,4,5-tetrahydropyridine-2,6-dicarboxylate N-succinyltransferase
EC 2.3.1.118 N-hydroxyarylamine O-acetyltransferase
EC 2.3.1.119 deleted, now covered by EC 2.3.1.199, EC 1.1.1.330 and EC 4.2.1.134
EC 2.3.1.120 deleted
EC 2.3.1.121 1-alkenylglycerophosphoethanolamine O-acyltransferase
EC 2.3.1.122 trehalose O-mycolyltransferase
EC 2.3.1.123 dolichol O-acyltransferase
EC 2.3.1.124 deleted
EC 2.3.1.125 1-alkyl-2-acetylglycerol O-acyltransferase
EC 2.3.1.126 isocitrate O-dihydroxycinnamoyltransferase
EC 2.3.1.127 ornithine N-benzoyltransferase
EC 2.3.1.128 transferred now EC 2.3.1.266 and EC 2.3.1.267
EC 2.3.1.129 acyl-[acyl-carrier-protein]UDP-N-acetylglucosamine O-acyltransferase
EC 2.3.1.130 galactarate O-hydroxycinnamoyltransferase
EC 2.3.1.131 glucarate O-hydroxycinnamoyltransferase
EC 2.3.1.132 glucarolactone O-hydroxycinnamoyltransferase
EC 2.3.1.133 shikimate O-hydroxycinnamoyltransferase
EC 2.3.1.134 galactolipid O-acyltransferase
EC 2.3.1.135 phosphatidylcholineretinol O-acyltransferase
EC 2.3.1.136 polysialic-acid O-acetyltransferase
EC 2.3.1.137 carnitine O-octanoyltransferase
EC 2.3.1.138 putrescine N-hydroxycinnamoyltransferase
EC 2.3.1.139 ecdysone O-acyltransferase
EC 2.3.1.140 rosmarinate synthase
EC 2.3.1.141 galactosylacylglycerol O-acyltransferase
EC 2.3.1.142 glycoprotein O-fatty-acyltransferase
EC 2.3.1.143 β-glucogallintetrakisgalloylglucose O-galloyltransferase
EC 2.3.1.144 anthranilate N-benzoyltransferase
EC 2.3.1.145 piperidine N-piperoyltransferase
EC 2.3.1.146 pinosylvin synthase
EC 2.3.1.147 glycerophospholipid arachidonoyl-transferase (CoA-independent)
EC 2.3.1.148 glycerophospholipid acyltransferase (CoA-dependent)
EC 2.3.1.149 platelet-activating factor acetyltransferase
EC 2.3.1.150 salutaridinol 7-O-acetyltransferase
EC 2.3.1.151 2,3',4,6-tetrahydroxybenzophenone synthase
EC 2.3.1.152 alcohol O-cinnamoyltransferase
EC 2.3.1.153 anthocyanin 5-(6'''-hydroxycinnamoyltransferase)
EC 2.3.1.154 transferred now EC 2.3.1.176
EC 2.3.1.155 acetyl-CoA C-myristoyltransferase
EC 2.3.1.156 phloroisovalerophenone synthase
EC 2.3.1.157 glucosamine-1-phosphate N-acetyltransferase
EC 2.3.1.158 phospholipid:diacylglycerol acyltransferase
EC 2.3.1.159 acridone synthase
EC 2.3.1.160 vinorine synthase
EC 2.3.1.161 lovastatin nonaketide synthase
EC 2.3.1.162 taxadien-5α-ol O-acetyltransferase
EC 2.3.1.163 10-hydroxytaxane O-acetyltransferase
EC 2.3.1.164 isopenicillin-N N-acyltransferase
EC 2.3.1.165 6-methylsalicylic acid synthase
EC 2.3.1.166 2α-hydroxytaxane 2-O-benzoyltransferase
EC 2.3.1.167 10-deacetylbaccatin III 10-O-acetyltransferase
EC 2.3.1.168 dihydrolipoyllysine-residue (2-methylpropanoyl)transferase
EC 2.3.1.169 CO-methylating acetyl-CoA synthase
EC 2.3.1.170 6'-deoxychalcone synthase
EC 2.3.1.171 anthocyanin 6"-O-malonyltransferase
EC 2.3.1.172 anthocyanin 5-O-glucoside 6'''-O-malonyltransferase
EC 2.3.1.173 flavonol-3-O-triglucoside O-coumaroyltransferase
EC 2.3.1.174 3-oxoadipyl-CoA thiolase
EC 2.3.1.175 deacetylcephalosporin-C acetyltransferase
EC 2.3.1.176 propanoyl-CoA C-acyltransferase
EC 2.3.1.177 3,5-dihydroxybiphenyl synthase
EC 2.3.1.178 diaminobutyrate acetyltransferase
EC 2.3.1.179 β-ketoacyl-[acyl-carrier-protein] synthase II
EC 2.3.1.180 β-ketoacyl-acyl-carrier-protein synthase III
EC 2.3.1.181 lipoyl(octanoyl) transferase
EC 2.3.1.182 (R)-citramalate synthase
EC 2.3.1.183 phosphinothricin acetyltransferase
EC 2.3.1.184 acyl-homoserine-lactone synthase
EC 2.3.1.185 tropine acyltransferase
EC 2.3.1.186 pseudotropine acyltransferase
EC 2.3.1.187 acetyl-S-ACP:malonate ACP transferase
EC 2.3.1.188 ω-hydroxypalmitate O-feruloyl transferase
EC 2.3.1.189 mycothiol synthase
EC 2.3.1.190 acetoin dehydrogenase system
EC 2.3.1.191 UDP-3-O-(3-hydroxymyristoyl)glucosamine N-acyltransferase
EC 2.3.1.192 glycine N-phenylacetyltransferase
EC 2.3.1.193 tRNAMet cytidine acetyltransferase
EC 2.3.1.194 acetoacetyl-CoA synthase
EC 2.3.1.195 (Z)-3-hexen-1-ol acetyltransferase
EC 2.3.1.196 benzyl alcohol O-benzoyltransferase
EC 2.3.1.197 dTDP-3-amino-3,6-dideoxy-α-D-galactopyranose 3-N-acetyltransferase
EC 2.3.1.198 glycerol-3-phosphate 2-O-acyltransferase
EC 2.3.1.199 very-long-chain 3-oxoacyl-CoA synthase
EC 2.3.1.200 lipoyl amidotransferase
EC 2.3.1.201 UDP-2-acetamido-3-amino-2,3-dideoxy-glucuronate N-acetyltransferase
EC 2.3.1.202 UDP-4-amino-4,6-dideoxy-N-acetyl-β-L-altrosamine N-acetyltransferase
EC 2.3.1.203 UDP-N-acetylbacillosamine N-acetyltransferase
EC 2.3.1.204 octanoyl-[GcvH]:protein N-octanoyltransferase
EC 2.3.1.205 fumigaclavine B O-acetyltransferase
EC 2.3.1.206 3,5,7-trioxododecanoyl-CoA synthase
EC 2.3.1.207 β-ketodecanoyl-[acyl-carrier-protein] synthase
EC 2.3.1.208 4-hydroxycoumarin synthase
EC 2.3.1.209 dTDP-4-amino-4,6-dideoxy-D-glucose acyltransferase
EC 2.3.1.210 dTDP-4-amino-4,6-dideoxy-D-galactose acyltransferase
EC 2.3.1.211 bisdemethoxycurcumin synthase
EC 2.3.1.212 benzalacetone synthase
EC 2.3.1.213 cyanidin 3-O-(6-O-glucosyl-2-O-xylosylgalactoside) 6'''-O-hydroxycinnamoyltransferase
EC 2.3.1.214 pelargonidin 3-O-(6-caffeoylglucoside) 5-O-(6-O-malonylglucoside) 4'''-malonyltransferase
EC 2.3.1.215 anthocyanidin 3-O-glucoside 6''-O-acyltransferase
EC 2.3.1.216 5,7-dihydroxy-2-methylchromone synthase
EC 2.3.1.217 curcumin synthase
EC 2.3.1.218 phenylpropanoylacetyl-CoA synthase
EC 2.3.1.219 demethoxycurcumin synthase
EC 2.3.1.220 2,4,6-trihydroxybenzophenone synthase
EC 2.3.1.221 noranthrone synthase
EC 2.3.1.222 phosphate propanoyltransferase
EC 2.3.1.223 3-oxo-5,6-didehydrosuberyl-CoA thiolase
EC 2.3.1.224 acetyl-CoA-benzylalcohol acetyltransferase
EC 2.3.1.225 protein S-acyltransferase
EC 2.3.1.226 carboxymethylproline synthase
EC 2.3.1.227 GDP-perosamine N-acetyltransferase
EC 2.3.1.228 isovaleryl-homoserine lactone synthase
EC 2.3.1.229 4-coumaroyl-homoserine lactone synthase
EC 2.3.1.230 2-heptyl-4(1H)-quinolone synthase
EC 2.3.1.231 tRNAPhe {7-[3-amino-3-(methoxycarbonyl)propyl]wyosine37-N}-methoxycarbonyltransferase
EC 2.3.1.232 methanol O-anthraniloyltransferase
EC 2.3.1.233 1,3,6,8-tetrahydroxynaphthalene synthase
EC 2.3.1.234 N6-L-threonylcarbamoyladenine synthase
EC 2.3.1.235 tetracenomycin F2 synthase
EC 2.3.1.236 5-methylnaphthoic acid synthase
EC 2.3.1.237 neocarzinostatin naphthoate synthase
EC 2.3.1.238 monacolin J acid methylbutanoate transferase
EC 2.3.1.239 10-deoxymethynolide synthase
EC 2.3.1.240 narbonolide synthase
EC 2.3.1.241 Kdo2-lipid IVA acyltransferase
EC 2.3.1.242 Kdo2-lipid IVA palmitoleoyltransferase
EC 2.3.1.243 acyl-Kdo2-lipid IVA acyltransferase
EC 2.3.1.244 2-methylbutanoate polyketide synthase
EC 2.3.1.245 3-hydroxy-5-phosphooxypentane-2,4-dione thiolase
EC 2.3.1.246 3,5-dihydroxyphenylacetyl-CoA synthase
EC 2.3.1.247 (5S)-5-amino-3-oxohexanoate:acetyl-CoA ethylamine transferase
EC 2.3.1.248 spermidine disinapoyl transferase
EC 2.3.1.249 spermidine dicoumaroyl transferase
EC 2.3.1.250 [Wnt protein] O-palmitoleoyl transferase
EC 2.3.1.251 lipid IVA palmitoyltransferase
EC 2.3.1.252 mycolipanoate synthase
EC 2.3.1.253 phloroglucinol synthase
EC 2.3.1.254 N-terminal methionine Nα-acetyltransferase NatB
EC 2.3.1.255 N-terminal amino-acid Nα-acetyltransferase NatA
EC 2.3.1.256 N-terminal methionine Nα-acetyltransferase NatC
EC 2.3.1.257 N-terminal L-serine Nα-acetyltransferase NatD
EC 2.3.1.258 N-terminal methionine Nα-acetyltransferase NatE
EC 2.3.1.259 N-terminal methionine Nα-acetyltransferase NatF
EC 2.3.1.260 tetracycline polyketide synthase
EC 2.3.1.261 4-hydroxyphenylalkanoate synthase
EC 2.3.1.262 anthraniloyl-CoA anthraniloyltransferase
EC 2.3.1.263 2-amino-4-oxopentanoate thiolase
EC 2.3.1.264 β-lysine N6-acetyltransferase
EC 2.3.1.265 phosphatidylinositol dimannoside acyltransferase
EC 2.3.1.266 [ribosomal protein bS18]-alanine N-acetyltransferase
EC 2.3.1.267 [ribosomal protein uS5]-alanine N-acetyltransferase
EC 2.3.1.268 ethanol O-acetyltransferase
EC 2.3.1.269 apolipoprotein N-acyltransferase
EC 2.3.1.270 lyso-ornithine lipid O-acyltransferase
EC 2.3.1.271 L-glutamate-5-semialdehyde N-acetyltransferase
EC 2.3.1.272 2-acetylphloroglucinol acetyltransferase
EC 2.3.1.273 diglucosylglycerate octanoyltransferase
EC 2.3.1.274 phosphate acyltransferase
EC 2.3.1.275 acyl phosphate:glycerol-3-phosphate acyltransferase
EC 2.3.1.276 galactosamine-1-phosphate N-acetyltransferase
EC 2.3.1.277 2-oxo-3-(phosphooxy)propyl 3-oxoalkanoate synthase
EC 2.3.1.278 mycolipenoyl-CoA—2-(long-chain-fatty acyl)-trehalose mycolipenoyltransferase
EC 2.3.1.279 long-chain-acyl-CoA—trehalose acyltransferase
EC 2.3.1.280 (aminoalkyl)phosphonate N-acetyltransferase
EC 2.3.1.281 5-hydroxydodecatetraenal polyketide synthase
EC 2.3.1.282 phenolphthiocerol/phthiocerol/phthiodiolone dimycocerosyl transferase
EC 2.3.1.283 2'-acyl-2-O-sulfo-trehalose (hydroxy)phthioceranyltransferase
EC 2.3.1.284 3'-(hydroxy)phthioceranyl-2'-palmitoyl(stearoyl)-2-O-sulfo-trehalose (hydroxy)phthioceranyltransferase
EC 2.3.1.285 (13S,14R)-1,13-dihydroxy-N-methylcanadine 13-O-acetyltransferase
EC 2.3.1.286 protein acetyllysine N-acetyltransferase
EC 2.3.1.287 phthioceranic/hydroxyphthioceranic acid synthase
EC 2.3.1.288 2-O-sulfo trehalose long-chain-acyltransferase
EC 2.3.1.289 aureothin polyketide synthase system
EC 2.3.1.290 spectinabilin polyketide synthase system
EC 2.3.1.291 sphingoid base N-palmitoyltransferase
EC 2.3.1.292 (phenol)carboxyphthiodiolenone synthase
EC 2.3.1.293 meromycolic acid 3-oxoacyl-(acyl carrier protein) synthase I
EC 2.3.1.294 meromycolic acid 3-oxoacyl-(acyl carrier protein) synthase II
EC 2.3.1.295 mycoketide-CoA synthase
EC 2.3.1.296 ω-hydroxyceramide transacylase
EC 2.3.1.297 very-long-chain ceramide synthase
EC 2.3.1.298 ultra-long-chain ceramide synthase
EC 2.3.1.299 sphingoid base N-stearoyltransferase
EC 2.3.1.300 branched-chain β-ketoacyl-[acyl-carrier-protein] synthase
EC 2.3.1.301 mycobacterial β-ketoacyl-[acyl carrier protein] synthase III
EC 2.3.1.302 hydroxycinnamoyl-CoA:5-hydroxyanthranilate N-hydroxycinnamoyltransferase
EC 2.3.1.303 α-L-Rha-(1→2)-α-D-Man-(1→2)-α-D-Man-(1→3)-α-D-Gal-PP-Und 2IV-O-acetyltransferase
EC 2.3.1.304 poly[(R)-3-hydroxyalkanoate] polymerase
EC 2.3.1.305 acyl-[acyl-carrier protein]—UDP-2-acetamido-3-amino-2,3-dideoxy-α-D-glucopyranose N-acyltransferase
EC 2.3.1.306 acetyl-CoA:lysine N6-acetyltransferase
EC 2.3.1.307 6-diazo-5-oxo-L-norleucine Nα-acetyltranferase
EC 2.3.1.308 tubulin N-terminal N-acetyltransferase NAT9
EC 2.3.1.309 [β-tubulin]-L-lysine N-acetyltransferase
EC 2.3.1.310 benzoylsuccinyl-CoA thiolase
EC 2.3.1.311 tRNA carboxymetyluridine synthase
EC 2.3.1.312 D-glutamate N-acetyltransferase
EC 2.3.1.313 NAD-dependent lipoamidase
EC 2.3.1.314 phytol O-acyltransferase
EC 2.3.1.315 succinyl-CoA:cyclohexane-1-carboxylate CoA transferase
EC 2.3.1.316 N-hydroxyputrescine acetyltransferase
EC 2.3.1.317 3-dehydrocarnitine:acetyl-CoA trimethylamine transferase
EC 2.3.1.318 3-oxoadipate:acetyl-CoA acetyltransferase
EC 2.3.1.319 3,5-dioxohexanoate:acetyl-CoA acetone transferase
EC 2.3.1.320 taxoid C-13 O-(3-amino-3-phenylpropanoyl)transferase
EC 2.3.1.321 3′-N-debenzoyltaxol N-benzoyltransferase
EC 2.3.1.322 akuammiline synthase
EC 2.3.1.323 stemmadenine O-acetyltransferase