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()-matrairesinol biosynthesis
EC 1.14.14.131 bursehernin 5'-monooxygenase
EC 1.14.19.72 ()-pluviatolide synthase
EC 1.14.20.8 ()-deoxypodophyllotoxin synthase
EC 2.1.1.323 ()-pluviatolide 4-O-methyltransferase
EC 2.1.1.330 5'-demethylyatein 5'-O-methyltransferase